Visualizing Dynamics of Wild Birds
The Irruptive Winter Migration of American Goldfinch
As described in Data Exploration Methods(details), partial
dependence functions (PDFs) summarize the effect of predictors on the
probability of occurrence after accounting for the average effect of
all other predictors. It is this "statistical control " that makes
partial dependence functions well suited to explore and visualize the
effects of predictors based on observational data.
In this section we use partial dependence functions to visualize the
dynamics of wild bird populations. We do this by computing the partial
dependence as a joint function of location and time, averaging out the
effects of all other predictors. The resulting estimates of occurrence
probability are plotted as surfaces and animated to visualize dynamic
patterns of occurrence.
The main advantage of these model-based partial dependence surface-plots over simpler surface-plots of “raw” observations is that they control for many important sources of variation that would otherwise bias distributional estimates. For example, the bagged decision tree models used to predict the probability of species’ occurrence take into account the effects of varying observer effort, factors that attract birds to feeders, and varying human population density. These effects are then “averaged out” by the partial dependence plots producing improved estimates of species’ occurrence. Another advantage of model-based partial dependence surface plots is that they can be used to estimate the potential for species’ occurrence at unobserved locations.
Using all of our PFW analyses for American Goldfinch we generated two types of animations to highlight distinct dynamic patterns of the winter migration. In Figure 3, we animate frames over each year at a common day within the Feederwatch season, December 1st. This animation allows one to view between-season differences in winter migration. From this animation the following patterns can be seen:
- Alternating high and low occurrence probability in the Ontario - Maine regions characterize the irruptive winter migration of American Goldfinch.
- There is a relatively stable region of high occurrence probability from South of the great lakes to the Central Hardwoods of the Ozark Mountains.
- Little change in the western population.

Figure 3. Animated Partial Dependence Surface-Plots for American Goldfinch showing the estimated probability of occurrence on February 15 from 1994 to 2004.
The second animation in Figure 4 allows one to visualize within-season
winter migration patterns. The animation below shows the estimated
probability of occurrence from November 15 – April 1 for the 2003-04
Feederwatch season. From this animation the following patterns can be
seen:
- The last part of the irruptive migration can be seen as the region
of high occurrence probability in the North East moves southward in
early winter.
- The region of high occurrence probability south of the great lakes
moves southward until February and then it appears to start moving
north as spring begins.

Figure 4. Animated Partial Dependence Surface-Plots for American Goldfinch showing the estimated probability of occurrence through the 2003-2004 winter season.